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Fat adds flavor and texture to baked goods:
http://busycooks.about.com/od/howtobake/a/bakingingredien.htm
http://www.ochef.com/225.htm
Will your cake still be edible, yes. Will it be a good as if you added the shortening, no.
But if you've never made or eaten this cake before, it could be passable...
HTH
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