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Well, the first reason that comes to mind is that the cake layers were over baked.
If you don't have a separate oven thermometer, I'd get one to make sure that your oven thermostat isn't off.
Additionally, I bake almost all of my cake layers at 325F and use both bake even strips and inverted flower nails (as needed).
Here's a handy list of cake trouble shooting charts:
http://tinyurl.com/2p5bdu
http://tinyurl.com/32goqe
http://tinyurl.com/6c745g
http://tinyurl.com/6lpjww
http://tinyurl.com/yay22w
HTH
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